An inconspicuous annual of open, often disturbed, habitats on chalk and heavy calcareous clay soils. It is found on grassy banks, verges, railway banks, woodland rides, and coastal grassland and shingle. Lowland.
There has been a considerable increase in 10-km square records since the 1962 Atlas, particularly in C.S. and S.E. England. Several recent county floras suggest that it is spreading in ruderal situations, and there are also records from restored open-cast mine sites where it may have been introduced as a constituent of commercial seed mixtures. It is unclear whether recent records in N.W. England are introductions or represent a northerly expansion of its native range.
European Temperate element.
PLANTATT - Attributes of British and Irish plants. (.zip 1455KB) This dataset was compiled and published in 2004, and last updated in November 2008. Download includes an Excel spreadsheet of the attributes, and a PDF explaining the background and nomenclature. Note that the PDF version is the booklet as published, whereas the Excel spreadsheet incorporates subsequent corrections. A hardcopy can be purchased from the Centre for Ecology and Hydrology.