Wilson, R.J., Davies, Z.G. & Thomas, C.D. (2009) Modelling the effect of habitat fragmentation on range expansion in a butterfly. Proceedings of the Royal Society B: Biological Sciences, 276, 1421-1427.
Botham, M.S., Ash, D., Aspey, N., Bourn, N. a. D., Bulman, C.R., Roy, D.B., Swain, J., Zannese, A. & Pywell, R.F. (2011) The effects of habitat fragmentation on niche requirements of the marsh fritillary, Euphydryas aurinia, (Rottemburg, 1775) on calcareous grasslands in southern UK. Journal of Insect Conservation, 15, 269-277.