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Botham, M.S., Ash, D., Aspey, N., Bourn, Na. D., Bulman, C.R., Roy, D.B., Swain, J., Zannese, A. & Pywell, R.F. (2011) The effects of habitat fragmentation on niche requirements of the marsh fritillary, Euphydryas aurinia, (Rottemburg, 1775) on calcareous grasslands in southern UK. Journal of Insect Conservation, 15, 269-277.

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Fox, R., Brereton, T.M., Asher, J., August, T.A., Botham, M.S., Bourn, Na. D., Cruickshanks, K.L., Bulman, C.R., Ellis, S., Harrower, C.A., Middlebrook, I., Noble, D.G., Powney, G.D., Randall, R.D., Warren, M.S. & Roy, D.B. (2015) The State Of The Uk’s Butterflies 2015. Butterfly Conservation and the Centre for Ecology & Hydrology, Wareham, Dorset.